The toes rule is mostly for D. tinctorius, D. azureus, D. leucomelas, and D. auratus. The males have much wider toe pads, but unless you are experienced in what normal male and female toepads look like, it won't be easy. I don't know how to do it. Usually you go by body shape, females are slightly larger, rounder, and males have bigger throat sacs(often hard to see) and as you said, call. Basically, if it calls, and it calls often and continuosly, its possibly a male. Until it lays eggs, its not one hundred percent certain that its a female.
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"Kings made tombs more splendid then the houses of the living. And counted the old names of their descent dearer than the names of their sons."
-J.R.R. Tolkien
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